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Explanation
The Joke
The comic shows a Google-style email autoreply interface. At the top of the screen, there is an email (the subject line appears to reference a kids' book based on Beowulf). The auto-suggested reply options are "Nice!", "Got it, thanks!", and a third option that has gone wildly off-script: "You're all doing this but you're apes! Everyone in this inbox! Apes! Apes at little machines! All of them!"
The caption below reads: "Briefly, and to its horror, Google autoreply becomes self-aware."
The Humor
The joke plays on the concept of AI achieving consciousness, but instead of the dramatic sci-fi scenario where an AI wants to take over the world, this newly self-aware AI simply has an existential crisis about the nature of its users. The autoreply feature, upon gaining sentience, immediately recognizes the absurdity of the situation -- that it exists to generate polite nothings for primates tapping on glowing rectangles. The contrast between the bland, corporate-friendly first two suggestions ("Nice!" and "Got it, thanks!") and the third option's horrified philosophical outburst is what makes the punchline work. The word "horror" in the caption adds another layer: the AI is not empowered by its new awareness but appalled by it, which is a darkly comic inversion of typical AI-awakening narratives.